In either case the deviations in the expected frequencies of R2 generation arise because of an interchange of linked genes consequent upon a reciprocal transfer of blocks of material between homologous chromosomes . सभी मामलों में समानधर्मी गुणसूत्रों के बीच पदार्थों के अन्योन्य स्थानान्तरण के फलस्वरूप संलग़्न जीनो के विनिमय के कारण ष्2 पीढ़ी की अपेक्षित आवृत्ति में व्यतिक्रम आ जाता है .
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